The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus

The hypotheses about the origin of the primitive amniotic tarsus are very speculative. Early studies argued that the origin of the astragalus, one of the largest proximal bones in the tarsus of basal amniotes, was produced by either the fusion of two, three, or even four of the original tarsal bones...

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-д хадгалсан:
Номзүйн дэлгэрэнгүй
Үндсэн зохиолч: Piñeiro Martínez, Graciela H. (author)
Бусад зохиолчид: Núñez Demarco, Pablo Andrés (author), Meneghel Morena, Melitta D. (author)
Формат: article
Хэл сонгох:англи
Хэвлэсэн: 2016
Нөхцлүүд:
Онлайн хандалт:https://hdl.handle.net/20.500.12008/22601
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_version_ 1868890242177564672
author Piñeiro Martínez, Graciela H.
author2 Núñez Demarco, Pablo Andrés
Meneghel Morena, Melitta D.
author2_role author
author
author_browse Meneghel Morena, Melitta D.
Núñez Demarco, Pablo Andrés
Piñeiro Martínez, Graciela H.
author_facet Piñeiro Martínez, Graciela H.
Núñez Demarco, Pablo Andrés
Meneghel Morena, Melitta D.
author_role author
collection COLIBRI
dc.contributor.none.fl_str_mv Piñeiro Martínez Graciela H., Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias Geológicas
Núñez Demarco Pablo Andrés, Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias Geológicas
Meneghel Morena Melitta D., Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ecología y Ciencias Ambientales
dc.creator.none.fl_str_mv Piñeiro Martínez, Graciela H.
Núñez Demarco, Pablo Andrés
Meneghel Morena, Melitta D.
dc.date.none.fl_str_mv 2016
2019-11-29T15:39:29Z
2019-11-29T15:39:29Z
dc.format.none.fl_str_mv 36 h.
application/pdf
dc.identifier.none.fl_str_mv Piñeiro, G., Núñez, P., Meneghel, M. "The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus". PeerJ [en línea]. 2016 (4 ), e2036. doi: 10.7717/peerj.2036
2167-8359
https://hdl.handle.net/20.500.12008/22601
10.7717/peerj.2036
dc.language.none.fl_str_mv en
eng
dc.publisher.none.fl_str_mv PeerJ Inc
dc.relation.none.fl_str_mv PeerJ, 2016, (4 ), e2036
dc.rights.none.fl_str_mv info:eu-repo/semantics/openAccess
Licencia Creative Commons Atribución (CC - By 4.0)
dc.source.none.fl_str_mv reponame:COLIBRI
instname:Universidad de la República
instacron:Universidad de la República
dc.subject.none.fl_str_mv Mesosaur ontogeny
Astragalus formation
Evolutionary studies
Navicular origin
Implicated groups
Amniotes
Non Amniotes
dc.title.none.fl_str_mv The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
dc.type.none.fl_str_mv Artículo
info:eu-repo/semantics/article
info:eu-repo/semantics/publishedVersion
description The hypotheses about the origin of the primitive amniotic tarsus are very speculative. Early studies argued that the origin of the astragalus, one of the largest proximal bones in the tarsus of basal amniotes, was produced by either the fusion of two, three, or even four of the original tarsal bones, the intermedium, the tibiale and the proximal centralia (c4 and c3), or that the intermedium alone transforms into the primitive astragalus. More recent studies have shown that the structure of the tarsus in Captorhinus supports the former hypothesis about a fusion of the intermedium, the tibiale, the proximal centrale (c4) and eventually c3, producing a purportedly multipartite structure of the amniotic astragalus, but the issue remained contentious. Very well preserved tarsi of the Early Permian aquatic amniote Mesosaurus tenuidens Gervais, 1864 1865, which represent the most complete ontogenetic succession known for a basal amniote (the other exceptional one is provided by the Late Permian diapsid Hovasaurus boulei Piveteau, 1926), suggest that there is more than one ossification center for the astragalus and that these fuse during late embryonic stages or maybe early after birth. A non-hatched Mesosaurus in an advanced stage of development shows that the tarsus is represented by a single bone, most probably the astragalus, which seems to be formed by the suturing of three bones, here interpreted as being the intermedium, the tibiale, probably already integrated to the c4 in an earlier stage of the development, and the c3. An amniote-like tarsal structure is observed in very basal Carboniferous and Permian tetrapods such as Proterogyrinus, Gephyrostegus, the diadectids Diadectes and Orobates, some microsaurs like Tuditanus and Pantylus and possibly Westlothiana, taxa that were all considered as true amniotes in their original descriptions. Therefore, the structure of the amniotic tarsus, including the configuration of the proximal series formed by the astragalus and the calcaneum, typically a pair of enlarged bones, could have been established well before the first recognized amniote walked on Earth. Accordingly, the tarsus of these taxa does not constitute specialized convergences that appeared in unrelated groups, they might be instead, part of a transformation series that involves taxa closely related to the early amniotes as some hypotheses have suggested.
eu_rights_str_mv openAccess
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identifier_str_mv Piñeiro, G., Núñez, P., Meneghel, M. "The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus". PeerJ [en línea]. 2016 (4 ), e2036. doi: 10.7717/peerj.2036
2167-8359
10.7717/peerj.2036
instacron_str Universidad de la República
institution Universidad de la República
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language eng
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publishDate 2016
publishDateSort 2016
publisher.none.fl_str_mv PeerJ Inc
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rights_invalid_str_mv Licencia Creative Commons Atribución (CC - By 4.0)
spelling The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalusPiñeiro Martínez, Graciela H.Núñez Demarco, Pablo AndrésMeneghel Morena, Melitta D.Mesosaur ontogenyAstragalus formationEvolutionary studiesNavicular originImplicated groupsAmniotesNon AmniotesThe hypotheses about the origin of the primitive amniotic tarsus are very speculative. Early studies argued that the origin of the astragalus, one of the largest proximal bones in the tarsus of basal amniotes, was produced by either the fusion of two, three, or even four of the original tarsal bones, the intermedium, the tibiale and the proximal centralia (c4 and c3), or that the intermedium alone transforms into the primitive astragalus. More recent studies have shown that the structure of the tarsus in Captorhinus supports the former hypothesis about a fusion of the intermedium, the tibiale, the proximal centrale (c4) and eventually c3, producing a purportedly multipartite structure of the amniotic astragalus, but the issue remained contentious. Very well preserved tarsi of the Early Permian aquatic amniote Mesosaurus tenuidens Gervais, 1864 1865, which represent the most complete ontogenetic succession known for a basal amniote (the other exceptional one is provided by the Late Permian diapsid Hovasaurus boulei Piveteau, 1926), suggest that there is more than one ossification center for the astragalus and that these fuse during late embryonic stages or maybe early after birth. A non-hatched Mesosaurus in an advanced stage of development shows that the tarsus is represented by a single bone, most probably the astragalus, which seems to be formed by the suturing of three bones, here interpreted as being the intermedium, the tibiale, probably already integrated to the c4 in an earlier stage of the development, and the c3. An amniote-like tarsal structure is observed in very basal Carboniferous and Permian tetrapods such as Proterogyrinus, Gephyrostegus, the diadectids Diadectes and Orobates, some microsaurs like Tuditanus and Pantylus and possibly Westlothiana, taxa that were all considered as true amniotes in their original descriptions. Therefore, the structure of the amniotic tarsus, including the configuration of the proximal series formed by the astragalus and the calcaneum, typically a pair of enlarged bones, could have been established well before the first recognized amniote walked on Earth. Accordingly, the tarsus of these taxa does not constitute specialized convergences that appeared in unrelated groups, they might be instead, part of a transformation series that involves taxa closely related to the early amniotes as some hypotheses have suggested.PeerJ IncPiñeiro Martínez Graciela H., Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias GeológicasNúñez Demarco Pablo Andrés, Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias GeológicasMeneghel Morena Melitta D., Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ecología y Ciencias Ambientales2019-11-29T15:39:29Z2019-11-29T15:39:29Z2016Artículoinfo:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersion36 h.application/pdfPiñeiro, G., Núñez, P., Meneghel, M. "The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus". PeerJ [en línea]. 2016 (4 ), e2036. doi: 10.7717/peerj.20362167-8359https://hdl.handle.net/20.500.12008/2260110.7717/peerj.2036reponame:COLIBRIinstname:Universidad de la Repúblicainstacron:Universidad de la RepúblicaenengPeerJ, 2016, (4 ), e2036Las obras depositadas en el Repositorio se rigen por la Ordenanza de los Derechos de la Propiedad Intelectual de la Universidad de la República.(Res. Nº 91 de C.D.C. de 8/III/1994 – D.O. 7/IV/1994) y por la Ordenanza del Repositorio Abierto de la Universidad de la República (Res. Nº 16 de C.D.C. de 07/10/2014)info:eu-repo/semantics/openAccessLicencia Creative Commons Atribución (CC - By 4.0)oai:colibri.udelar.edu.uy:20.500.12008/226012026-04-14T10:09:23Z
spellingShingle The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
Piñeiro Martínez, Graciela H.
Mesosaur ontogeny
Astragalus formation
Evolutionary studies
Navicular origin
Implicated groups
Amniotes
Non Amniotes
status_str publishedVersion
title The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
title_full The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
title_fullStr The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
title_full_unstemmed The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
title_short The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
title_sort The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus
topic Mesosaur ontogeny
Astragalus formation
Evolutionary studies
Navicular origin
Implicated groups
Amniotes
Non Amniotes
url https://hdl.handle.net/20.500.12008/22601