HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo

Ensifer meliloti is a nitrogen-fixing symbiont of the alfalfa legume able to use heme as an iron source. The transport mechanism involved in heme acquisition in E. meliloti has been identified and characterized, but the fate of heme once inside the cell is not known. In silico analysis of E. melilot...

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1. Verfasser: Amarelle, Vanesa (author)
Weitere Verfasser: Rosconi, Federico (author), Lázaro-Mártinez, Juan Manuel (author), Buldain, Graciela (author), Noya, Francisco (author), O´Brian, Mark R. (author), Fabiano, Elena (author)
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Sprache:Englisch
Veröffentlicht: 2016
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Online-Zugang:http://hdl.handle.net/20.500.12381/200
http://dx.doi.org/10.1007/s10534-016-9919-3
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author Amarelle, Vanesa
author2 Rosconi, Federico
Lázaro-Mártinez, Juan Manuel
Buldain, Graciela
Noya, Francisco
O´Brian, Mark R.
Fabiano, Elena
author2_role author
author
author
author
author
author
author_browse Amarelle, Vanesa
Buldain, Graciela
Fabiano, Elena
Lázaro-Mártinez, Juan Manuel
Noya, Francisco
O´Brian, Mark R.
Rosconi, Federico
author_facet Amarelle, Vanesa
Rosconi, Federico
Lázaro-Mártinez, Juan Manuel
Buldain, Graciela
Noya, Francisco
O´Brian, Mark R.
Fabiano, Elena
author_role author
collection REDI
dc.creator.none.fl_str_mv Amarelle, Vanesa
Rosconi, Federico
Lázaro-Mártinez, Juan Manuel
Buldain, Graciela
Noya, Francisco
O´Brian, Mark R.
Fabiano, Elena
dc.date.none.fl_str_mv 2016
2019-11-05T20:42:32Z
2019-11-05T20:42:32Z
dc.format.none.fl_str_mv 17 p.
dc.identifier.none.fl_str_mv http://hdl.handle.net/20.500.12381/200
PR_FCE_2009_1_2564
http://dx.doi.org/10.1007/s10534-016-9919-3
dc.language.none.fl_str_mv eng
dc.publisher.none.fl_str_mv Springer
dc.rights.none.fl_str_mv Acceso abierto
info:eu-repo/semantics/openAccess
Reconocimiento-NoComercial-SinObraDerivada. (CC BY-NC-ND)
dc.source.none.fl_str_mv BioMetals. 2016; 29(2)
reponame:REDI
instname:Agencia Nacional de Investigación e Innovación
instacron:Agencia Nacional de Investigación e Innovación
dc.subject.none.fl_str_mv Rizobios
Hemo-oxigenasas
Metabolismo de hierro
Degradación de hemo
Ciencias Naturales y Exactas
Ciencias Biológicas
Bioquímica y Biología Molecular
dc.title.none.fl_str_mv HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
dc.type.none.fl_str_mv Artículo
info:eu-repo/semantics/article
Publicado
info:eu-repo/semantics/publishedVersion
description Ensifer meliloti is a nitrogen-fixing symbiont of the alfalfa legume able to use heme as an iron source. The transport mechanism involved in heme acquisition in E. meliloti has been identified and characterized, but the fate of heme once inside the cell is not known. In silico analysis of E. meliloti 1021 genome revealed no canonical heme oxygenases although two genes encoding putative heme degrading enzymes, smc01518 and hmuS, were identified. SMc01518 is similar to HmuQ of Bradyrhizobium japonicum, which is weakly homologous to the Staphylococcus aureus IsdG heme-degrading monooxygenase, whereas HmuS is homolog to Pseudomonas aeruginosa PhuS, a protein reported as a heme chaperone and as a heme degrading enzyme. Recombinant HmuQ and HmuS were able to bind hemin with a 1:1 stoichiometry and displayed a Kd value of 5 and 4 lM, respectively. HmuS degrades heme in vitro to the biliverdin isomers IX-b and IX-d in an equimolar ratio. The HmuQ recombinant protein degrades heme to biliverdin IX-d only. Additionally, in this work we demonstrate that humS and hmuQ gene expression is regulated by iron and heme in a RirA dependent manner and that both proteins are involved in heme metabolism in E. meliloti in vivo.
eu_rights_str_mv openAccess
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id anni_8e64bc31ba7b74deb6ec10ef2a7b6de9
identifier_str_mv PR_FCE_2009_1_2564
instacron_str Agencia Nacional de Investigación e Innovación
institution Agencia Nacional de Investigación e Innovación
instname_str Agencia Nacional de Investigación e Innovación
language eng
network_acronym_str anni
network_name_str oai-lr-anni
oai_identifier_str oai:redi.anii.org.uy:20.500.12381/200
publishDate 2016
publishDateSort 2016
publisher.none.fl_str_mv Springer
reponame_str REDI
repository.mail.fl_str_mv
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rights_invalid_str_mv Acceso abierto
Reconocimiento-NoComercial-SinObraDerivada. (CC BY-NC-ND)
spelling HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivoAmarelle, VanesaRosconi, FedericoLázaro-Mártinez, Juan ManuelBuldain, GracielaNoya, FranciscoO´Brian, Mark R.Fabiano, ElenaRizobiosHemo-oxigenasasMetabolismo de hierroDegradación de hemoCiencias Naturales y ExactasCiencias BiológicasBioquímica y Biología MolecularEnsifer meliloti is a nitrogen-fixing symbiont of the alfalfa legume able to use heme as an iron source. The transport mechanism involved in heme acquisition in E. meliloti has been identified and characterized, but the fate of heme once inside the cell is not known. In silico analysis of E. meliloti 1021 genome revealed no canonical heme oxygenases although two genes encoding putative heme degrading enzymes, smc01518 and hmuS, were identified. SMc01518 is similar to HmuQ of Bradyrhizobium japonicum, which is weakly homologous to the Staphylococcus aureus IsdG heme-degrading monooxygenase, whereas HmuS is homolog to Pseudomonas aeruginosa PhuS, a protein reported as a heme chaperone and as a heme degrading enzyme. Recombinant HmuQ and HmuS were able to bind hemin with a 1:1 stoichiometry and displayed a Kd value of 5 and 4 lM, respectively. HmuS degrades heme in vitro to the biliverdin isomers IX-b and IX-d in an equimolar ratio. The HmuQ recombinant protein degrades heme to biliverdin IX-d only. Additionally, in this work we demonstrate that humS and hmuQ gene expression is regulated by iron and heme in a RirA dependent manner and that both proteins are involved in heme metabolism in E. meliloti in vivo.Agencia Nacional de Investigación e InnovaciónSpringer2019-11-05T20:42:32Z2019-11-05T20:42:32Z2016Artículoinfo:eu-repo/semantics/articlePublicadoinfo:eu-repo/semantics/publishedVersion17 p.http://hdl.handle.net/20.500.12381/200PR_FCE_2009_1_2564http://dx.doi.org/10.1007/s10534-016-9919-3BioMetals. 2016; 29(2)reponame:REDIinstname:Agencia Nacional de Investigación e Innovacióninstacron:Agencia Nacional de Investigación e InnovaciónengAcceso abiertoinfo:eu-repo/semantics/openAccessReconocimiento-NoComercial-SinObraDerivada. (CC BY-NC-ND)oai:redi.anii.org.uy:20.500.12381/2002026-06-16T05:02:45Z
spellingShingle HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
Amarelle, Vanesa
Rizobios
Hemo-oxigenasas
Metabolismo de hierro
Degradación de hemo
Ciencias Naturales y Exactas
Ciencias Biológicas
Bioquímica y Biología Molecular
status_str publishedVersion
title HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
title_full HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
title_fullStr HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
title_full_unstemmed HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
title_short HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
title_sort HmuS and HmuQ of Ensifer/Sinorhizobium meliloti degrade heme in vitro and participate in heme metabolism in vivo
topic Rizobios
Hemo-oxigenasas
Metabolismo de hierro
Degradación de hemo
Ciencias Naturales y Exactas
Ciencias Biológicas
Bioquímica y Biología Molecular
url http://hdl.handle.net/20.500.12381/200
http://dx.doi.org/10.1007/s10534-016-9919-3